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    Echinodorus berteroi (Spreng.) Fassett

    Echinodorus berteroi (Spreng.) Fassett



    Alisma berteroanum Balb. ex Schult. & Schult.f.

    Alisma berteroi Spreng.

    Alisma macrophyllum var. minus Seub.

    Alisma rostratum Nutt.

    Alisma sprengelii Rich. ex Kunth

    Echinodorus berteroi subsp. patagonicus (Speg.) Rataj

    Echinodorus berteroi var. berteroi

    Echinodorus berteroi var. lanceolatus (Engelm. ex S.Watson & Coult.) Fernald

    Echinodorus berteroi var. patagonicus (Speg.) Rataj

    Echinodorus cordifolius f. lanceolatus (Engelm. ex S.Watson & Coult.) Fernald

    Echinodorus cordifolius var. berteroanus (Balb. ex Schult. & Schult.f.) Griseb.

    Echinodorus cordifolius var. lanceolatus (Engelm. ex S.Watson & Coult.) Mack. & Bush

    Echinodorus longilineatus Rataj

    Echinodorus patagonicus Speg.

    Echinodorus radicans (Nutt.) Engelm. ex A. Gray

    Echinodorus rostratus (Nutt.) Engelm.

    Echinodorus rostratus f. lanceolatus (Engelm. ex S.Watson & Coult.) Fernald

    Echinodorus rostratus var. lanceolatus Engelm. ex S.Watson & Coult.

    Sagittaria rostrata (Nutt.) Kuntze


    USA, Caribbean islands, Mexico, Pacific coast of Ecuador and Peru, Argentine and Paraguayan Chaco and Patagonia.


    Temporal pools and tidal flats. From sea level to 1500 m.

    Morphology General habit.

    Annual, glabrous, up to 110 cm.

    Morphology Leaves

    Leaves emersed or submerged, emerse blade elliptic-lanceolate, 3 - 11-veined, 3 - 15 cm long, 1 - 19 cm wide, the translucent markings present as strokes, apex acute to round, base obtuse to cordate, petioles round to triangular in cross-section, angled, 3 - 55 cm long, 5 mm diam, base with sheath up to 7 cm long, submerged blades linear, with 1 - 3 veins, ca. 15 cm long, 1 - 1.5 cm wide, the transparent markings absent or present as lines, the apex acute, the base attenuate, petioles triangular in cross-section, up to 4 cm long, 1 mm in diam., the base with a sheath up to 3cm long

    Morphology Reproductive morphology Fruits

    Fruit oblique, 4 - 6-ribbed, 2-keeled, glandular, 1.5 - 3 mm long, 0.5 - 1 mm wide, glands 1 - 2, separated by ribs, elliptic, beak end, erect, 0.9 - 1.8 mm.

    Morphology Reproductive morphology Inflorescences

    Inflorescence racemose or paniculate, of 1 - 9 whorls, each 3 - 18-flowered, erect, leaves overlapping, non-proliferating, up to 50 cm long, up to 25 cm wide, rachis triangular in cross-section, pedicels round, angled, 7 - 60 cm long, 5 mm diam, bracts free, lanceolate, 0.5 - 3.5 cm long, 2 - 7 mm wide, 5 - 10 veins, apex acute, pedicels in flowers and fruits spreading, round, 1 - 3 cm long, 0.3 mm diam. flowers 1 - 1.5 cm in diameter, sepals and corollas spreading, sepals 10 - 13-veined, ca. 5 mm long, c. 3 mm wide, veins without papillae, petals white, with claws, not overlapping, c. 7 mm long, c. 6 mm wide, stamens 13 - 15, the anthers many-sided, c. 0.5 mm long, the filaments c. 2 mm long, carpels numerous


    This species is resolved as sister to the rest of the genus in morphology-based (Lehtonen 2006), DNA-based, and complete evidence analysis (Lehtonen & Myllys 2008), but nuclear DNA indicated a drastically different phylogenetic position for the species. Since Echinodorus berteroi is the type species of the genus, its phylogenetic position determines the generic name of the other species discussed here. Should more comprehensive studies in the future place E. berteroi in a non-monophyletic position with the other species discussed here, a new genus should be established for the remaining species. South American populations of E. berteroi are morphologically similar to those from North America, and proposed species (E. patagonicus, Spegazzini 1902; E. longilineatus, Rataj 2004) or subspecific (E. berteroi subsp. patagonicus, Rataj 2004) rankings for South American populations are not warranted.


    Flowering and fruiting throughout the year in South America, from April to October in the United States.[1]


    Zuerst veröffentlicht in Rhodora 57: 139 (1955)

    Akzeptiert von:

    • Acevedo-Rodríguez, P. & Strong, M.T. (2005). Monocotyledons and Gymnosperms of Puerto Rico and the Virgin Islands Contributions from the United States National Herbarium 52: 1-415.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Adams, C.D. (1972). Flowering Plants of Jamaica: 1-848. University of the West Indies, Mona.
    • Allred, K.W. (2012). Flora Neomexicana, ed. 2, 1: 1-599. Range Science Herbarium, Las Cruces, New Mexico.
    • Bailey, C. & al. (2015). Guide to the Vascular Plants of Tennessee: 1-813. University of Tennessee press.
    • Boggan, J. Funck, V. & Kelloff, C. (1997). Checklist of the Plants of the Guianas (Guyana, Surinam, Franch Guiana) ed. 2: 1-238. University of Guyana, Georgetown.
    • Correll, D.S. & Correll, H.B. (1982). Flora of the Bahama Archipelago: 1-1692. J.Cramer, Vaduz.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Espejo Serena, A. & López-Ferrari, A.R. (1993). Las Monocotiledóneas Mexicanas una Sinopsis Florística 1(1): 1-76. Consejo Nacional de la Flora de México, México D.F.
    • Flora of North America Editorial Committee (2000). Flora of North America North of Mexico 22: 1-352. Oxford University Press, New York, Oxford.
    • Fournet, J. (2002). Flore illustrée des phanérogames de Guadeloupe et de Martinique 2: 1325-2538. Gondwana editions.
    • Govaerts, R. (2001). World Checklist of Seed Plants Database in ACCESS E-F: 1-50919.
    • Jones, R.L. (2005). Plant life of Kentucky. An illustrated guide to the vascular flora: 1-833. The universitry press of Kentucky.
    • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741.
    • Lehtonen, S. & Gordon, E. (2010). Actualización del conocimiento de los géneros Echinodorus y Helanthium (Alismataceae) en Venezuela Acta Botánica Venezuelica 33: 249-272.
    • Lehtonen, S. (2008 publ. 2009). An integative approach to species delimination in Echinodorus (Alismataceae) and the description of two new species Kew Bulletin 63: 525-563.
    • Liogier, H.A. & Martorell, L.F. (2000). Flora of Puerto Rico and Adjacent Islands: a Systematic Synopsis ed. 2: 1-382. Editorial de la Universidad de Puerto Rico, San Juan.
    • Zuloaga, F.O. & Morrone, O. (eds.) (1996). Pteridophyta, Gymnospermae y Angiospermae (Monocotyledoneae) Catálogo de las Plantas Vasculares de la República Argentina 1: 1-323. Missouri Botanical Garden, St. Louis.[6]

    1. POWO (2021). "Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; Retrieved 23 August 2021."
    2. 2.0 2.1 2.2 2.3 Roman Trifonov, Moskau Aquarium Pflanzen
    3. 3.0 3.1 3.2 (c) Nathan Taylor, einige Rechte vorbehalten (CC BY-NC),
    4. Zoya Akulova
    5. U.S. Army Corps of Engineers Foto von George Delancey
    6. Kew Names and Taxonomic Backbone The International Plant Names Index and World Checklist of Selected Plant Families 2021. Published on the Internet at and © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families.