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    Aquarius pubescens (Mart. ex Schult.f.) Christenh. & Byng

    Aquarius pubescens (Mart. ex Schult.f.) Christenh. & Byng


    Basionym : Echinodorus pubescens (Mart. ex Schult.f.) Seub. ex Warm.[1]


       Echinodorus ellipticus var. pubescens (Mart. ex Schult.f.) Micheli

       Echinodorus macrocarpus Rataj

       Echinodorus palifolius var. pubescens (Mart. ex Schult.f.) J.F.Macbr.


    Eastern Brazil.


    From sea level to 500 m.

    Morphology General habit

    Perennial, from rhizomes, petioles and stems hairy with simple hairs, up to 120 cm, rhizomes 3 cm diam. Leaves emersed, leaf blade ovate to elliptic, 5 - 9-veined, 10 - 24 cm long, 3 - 11 cm wide, translucent markings absent or present as lines, apex round to acute, base thin to cordate, petioles round, 3 - 35 cm long, c. 5 mm diam., base with sheath up to 10 cm long

    Morphology Reproductive morphology Fruits

    Fruit oblanceoloid, c. 7-ribbed, glandular, 2.5 - 3.5 mm long, 1.6 mm wide, glands 1 - 3, often severed from the ribs, elliptic, beak end, erect, c. 1 mm.

    Morphology Reproductive morphology Inflorescences

    Inflorescence racemose or paniculate, of 6 - 18 whorls, each 9 - 15-flowered, erect, overhanging leaves, non-propagative, 20 - 70 cm long, 15 cm wide, leaf blade triangular in cross-section, petioles round, up to 55 cm long , C. 5 mm in diameter, bracts fused at the base, lanceolate, coarse with membranous margins, up to 2 cm long, 5 mm wide, 11 - 16-veined, apex pointed, pedicels spreading, spreading in flowers and fruits, round, 0.5 - 1 cm long , 0.5 mm diam. flowers 1.5 - 2 cm in diameter, sepals erect, 16 - 21-veined, ca. 5 mm long, ca. 4 mm wide, the veins pubescent, petals spreading, white, not clawed, not overlapping, ca. 15 mm long, ca. 14 mm wide, stamens 12, anthers many-sided, ca. 1.5 mm long, filaments c. 1.5 mm long, carpels numerous

    Echinodorus macrocarpus was synonymized with E. paniculatus by Haynes & Holm-Nielsen (1994). However, the type specimen (Eugenio 227 RB!) has none of the typical characteristics of E. paniculatus. Instead, E. macrocarpus is here considered specific to E. pubescens due to a number of typical features of E. pubescens: ovate leaves without pellucid markings, large, densely ribbed achenes with 2 - 3 glands and stems covered with simple hairs.


    Flowering and fruiting from January to July.[1]

    1. 1.0 1.1 POWO (2021). "Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Im Internet veröffentlicht; Abgerufen am 22. August 2021."
    2. 2.0 2.1 2.2 y The New York Botanical Garden (licensed under
    3. Staatliche Naturwissenschaftliche Sammlungen Bayerns. The Vascular Plant Collection at the Botanische Staatssammlung München. Occurrence dataset
    4. by Naturalis Biodiversity Center (licensed under